B.A., Physics, University of Pennsylvania; Ph.D., Biology, California Institute of Technology (Caltech); post-doc, University of California Santa Cruz, Biology; visiting scientist, Marine Biological Laboratory & Woods Hole Oceanographic Institute, Woods Hole, MA, USA; rose through the professorial ranks at Wayne State University, Detroit, MI, USA. Ram's research on marine worms in Wales, England, and Woods Hole led to discoveries about glutathione that ultimately resulted in research of its effects on the extracellular calcium sensing receptor.